Debuysschere, M., Gheerbrant, E. & Allain, R. Earliest known European mammals: a review of the Morganucodonta from Saint-Nicolas-de-Port (Upper Triassic, France). J. Syst. Paleontol. 13, 825–855 (2015).
Google Scholar
Irmis, R., Nesbitt, S. J. & Sues, H.-D. in Anatomy, Phylogeny and Palaeobiology of Early Archosaurs and Their Kin Vol. 239 (eds Nesbitt, S. J. et al.) 275–302 (2013).
Jones, M. E. H. et al. Integration of molecules and new fossils supports a Triassic origin for Lepidosauria (lizards, snakes, and tuatara). BMC Evol. Biol. 13, 208 (2013).
Google Scholar
Langer, M. C., Ramezani, J. & Da Rosa, Á. A. S. U–Pb age constraints on dinosaur rise from south Brazil. Gondwana Res. 57, 133–140 (2018).
Google Scholar
Li, C., Wu, X.-C., Rieppel, O., Wang, L.-T. & Zhao, L.-J. An ancestral turtle from the Late Triassic of southwestern China. Nature 456, 497–501 (2008).
Google Scholar
Lyson, T. R. & Bever, G. S. Origin and evolution of the turtle body plan. Ann. Rev. Ecol. Evol. Syst. 51, 143–166 (2020).
Google Scholar
Mao, F., Zhang, C., Liu, C. & Meng, J. Fossoriality and evolutionary development in two Cretaceous mammaliamorphs. Nature 592, 577–582 (2021).
Google Scholar
Schoch, R. R., Werneburg, R. & Voigt, S. A Triassic stem-salamander from Kyrgyzstan and the origin of salamanders. Proc. Natl Acad. Sci. USA 117, 11584–11588 (2020).
Google Scholar
Simões, T. R. et al. The origin of squamates revealed by a Middle Triassic lizard from the Italian Alps. Nature 557, 706–709 (2018).
Google Scholar
Stocker, M. R. et al. The earliest equatorial record of frogs from the Late Triassic of Arizona. Biol. Lett. 15, 20180922 (2019).
Google Scholar
Martínez, R. N., Simões, T. R., Sobral, G. & Apesteguía, S. A Triassic stem lepidosaur illuminates the origin of lizard-like reptiles. Nature 597, 235–238 (2021).
Button, D. J., Lloyd, G. T., Ezcurra, M. D. & Butler, R. J. Mass extinctions drove increased global faunal cosmopolitanism on the supercontinent Pangaea. Nat. Commun. 8, 733 (2017).
Google Scholar
Chen, Z.-Q. & Benton, M. J. The timing and pattern of biotic recovery following the end-Permian mass extinction. Nat. Geosci. 5, 375–383 (2012).
Google Scholar
Roopnarine, P. D., Angielczyk, K. D., Wang, S. C. & Hertog, R. Trophic network models Eeplain instability of Early Triassic terrestrial communities. Proc. Biol. Sci. 274, 2077–2086 (2007).
Google Scholar
Ezcurra, M. D. Biogeography of Triassic tetrapods: evidence for provincialism and driven sympatric cladogenesis in the early evolution of modern tetrapod lineages. Proc. R. Soc. B 277, 2547–2552 (2010).
Google Scholar
Kent, D. V. & Clemmensen, L. B. Northward dispersal of dinosaurs from Gondwana to Greenland at the mid-Norian (215–212 Ma, Late Triassic) dip in atmospheric pCO2. Proc. Natl Acad. Sci. USA 118, e2020778118 (2021).
Google Scholar
Whiteside, J. H., Grogan, D. S., Olsen, P. E. & Kent, D. V. Climatically driven biogeographic provinces of Late Triassic tropical Pangea. Proc. Natl Acad. Sci. USA 108, 8972–8977 (2011).
Google Scholar
Whiteside, J. H. et al. Extreme ecosystem instability suppressed tropical dinosaur dominance for 30 million years. Proc. Natl Acad. Sci. USA 112, 7909–7913 (2015).
Google Scholar
Langer, M. C. Studies on continental Late Triassic tetrapod biochronology. II. The Ischigualastian and a Carnian global correlation. J. South Amer. Earth Sci. 19, 219–239 (2005).
Google Scholar
Martínez, R. N. et al. Vertebrate succession in the Ischigualasto Formation. J. Vertebr. Paleontol. 32, 10–30 (2012).
Google Scholar
Colombi, C. et al. A high-precision U–Pb zircon age constraints the timing of the faunistic and palynofloristic events of the Carnian Ischigualasto Formation, San Juan, Argentina. J. South Amer. Earth Sci. 111, 103433 (2021).
Google Scholar
Novas, F. E., Ezcurra, M. D., Chatterjee, S. & Kutty, T. S. New dinosaur species from the Upper Triassic Upper Maleri and Lower Dharmaram formations of Central India. Earth Environ. Sci. Trans. R. Soc. Edin. 101, 333–349 (2010).
Sellwood, B. W. & Valdes, P. J. Mesozoic climates: general circulation models and the rock record. Sediment Geol. 190, 269–287 (2006).
Google Scholar
Kligman, B. T., Marsh, A. D., Sues, H.-D. & Sidor, C. A. A new non-mammalian eucynodont from the Chinle Formation (Triassic: Norian), and implications for the early Mesozoic equatorial cynodont record. Biol. Lett. 16, 20200631 (2020).
Google Scholar
Irmis, R. B. Evaluating hypotheses for the early diversification of dinosaurs. Earth Environ. Sci. Trans. R. Soc. Edin. 101, 397–426 (2010).
Cabreira, S. F. et al. A unique Late Triassic dinosauromorph assemblage reveals dinosaur ancestral anatomy and diet. Curr. Biol. 26, 3090–3095 (2016).
Google Scholar
Irmis, R. B., Mundil, R., Martz, J. W. & Parker, W. G. High-resolution U–Pb ages from the Upper Triassic Chinle Formation (New Mexico, USA) support a diachronous rise of dinosaurs. Earth Planet. Sci. Lett. 309, 258–267 (2011).
Google Scholar
Langer, M. C., Ezcurra, M. D., Bittencourt, J. S. & Novas, F. E. The origin and early evolution of dinosaurs. Biol. Rev. 85, 55–110 (2010).
Google Scholar
Raath, M. A., Oesterlen, P. M. & Kitching, J. W. First record of Triassic Rhynchosauria (Reptilia: Diapsida) from the Lower Zambezi Valley, Zimbabwe. Palaeontologia Africana 29, 1–10 (1992).
Oesterlen, P. M. The geology of the Dande West Area, Lower Zambezi Valley. Zimbabwe Geol. Soc. Bull. 98, 1–85 (1998).
Sciscio, L. et al. Sedimentology and palaeontology of the Upper Karoo Group in the Mid-Zambezi Basin, Zimbabwe: new localities and their implications for interbasinal correlation. Geol. Mag. 158, 1035–1058 (2021).
Griffin, C. T. Developmental patterns and variation among early theropods. J. Anat. 232, 604–640 (2018).
Google Scholar
Griffin, C. T. & Nesbitt, S. J. Anomalously high variation in postnatal development is ancestral for dinosaurs but lost in birds. Proc. Natl Acad. Sci. USA 113, 14757–14762 (2016).
Google Scholar
Griffin, C. T. et al. Assessing ontogenetic maturity in extinct saurian reptiles. Biol. Rev. 96, 470–525 (2021).
Google Scholar
Prieto-Márquez, A. & Norell, M. A. Redescrip tion of a nearly complete skull of Plateosaurus (Dinosauria: Sauropodomorpha) from the Late Triassic of Trossingen (Germany). in American Museum Novitates 1–58 (American Museum of Natural History, 2011).
Yates, A. M. The species taxonomy of the sauropodomorph dinosaurs from the Löwenstein Formation (Norian, Late Triassic) of Germany. Palaeontology 46, 317–337 (2003).
Google Scholar
Pretto, F. A., Langer, M. C. & Schultz, C. L. A new dinosaur (Saurischia: Sauropodomorpha) from the Late Triassic of Brazil provides insights on the evolution of sauropodomorph body plan. Zool. J. Linnean Soc. 185, 388–416 (2018).
Google Scholar
Ezcurra, M. D. A new early dinosaur (Saurischia: Sauropodomorpha) from the Late Triassic of Argentina: a reassessment of dinosaur origin and phylogeny. J. Syst. Paleontol. 8, 371–425 (2010).
Google Scholar
Baron, M. G., Norman, D. B. & Barrett, P. M. A new hypothesis of dinosaur relationships and early dinosaur evolution. Nature 543, 501–506 (2017).
Google Scholar
Nesbitt, S. J. The early evolution of archosaurs: relationships and the origin of major clades. in Bulletin of the American Museum Natural History 1–292 (American Museum of Natural History, 2011).
Langer, M. C., McPhee, B. W., Marsola, J. C. D. A., Roberto-da-Silva, L. & Cabreira, S. F. Anatomy of the dinosaur Pampadromaeus barberenai (Saurischia—Sauropodomorpha) from the Late Triassic Santa Maria Formation of southern Brazil. PLoS ONE 14, e0212543 (2019).
Google Scholar
Martinez, R. N. & Alcober, O. A. A basal sauropodomorph (Dinosauria: Saurischia) from the Ischigualasto Formation (Triassic, Carnian) and the early evolution of Sauropodomorpha. PLoS ONE 4, e4397 (2009).
Google Scholar
Sereno, P. C., Martínez, R. N. & Alcober, O. A. Osteology of Eoraptor lunensis (Dinosauria, Sauropodomorpha). J. Vertebr. Paleontol. 32, 83–179 (2012).
Google Scholar
Stefanic, C. M. & Nesbitt, S. J. The evolution and role of the hyposphene–hypantrum articulation in Archosauria: phylogeny, size and/or mechanics? R. Soc. Open Sci. 6, 190258 (2019).
Google Scholar
Marsola, J. C. A. et al. A new dinosaur with theropod affinities from the Late Triassic Santa Maria Formation, south Brazil. J. Vertebr. Paleontol. 38, e1531878 (2019).
Google Scholar
Langer, M. C. The pelvic and hind limb anatomy of the stem-sauropodomorph Saturnalia tupiniquim (Late Triassic, Brazil). PaleoBios 23, 1–30 (2003).
Moro, D., Kerber, L., Müller, R. T. & Pretto, F. A. Sacral co-ossification in dinosaurs: the oldest record of fused sacral vertebrae in Dinosauria and the diversity of sacral co-ossification patterns in the group. J. Anat. 238, 828–844 (2021).
Google Scholar
Sereno, P. C. The pectoral girdle and forelimb of the basal theropod Herrerasaurus ischigualastensis. J. Vertebr. Paleontol. 13, 425–450 (1994).
Google Scholar
Langer, M. C., França, M. A. G. & Gabriel, S. The pectoral girdle and forelimb anatomy of the stem-sauropodomorph Saturnalia tupiniquim (Upper Triassic, Brazil). Spec. Papers Palaeontol. 77, 113–137 (2007).
Langer, M. C., Bittencourt, J. S. & Schultz, C. L. A reassessment of the basal dinosaur Guaibasaurus candelariensis, from the Late Triassic Caturrita Formation of south Brazil. Earth Environ. Sci. Trans. R. Soc. Edin. 101, 301–332 (2010).
Yates, A. M. A new species of the primitive dinosaur Thecodontosaurus (Saurischia: Sauropodomorpha) and its implications for the systematics of early dinosaurs. J. Syst. Paleontol. 1, 1–42 (2003).
Google Scholar
Novas, F. E. New information on the systematics and postcranial skeleton of Herrerasaurus ischigualastensis (Theropoda: Herrerasauridae) from the Ischigualasto Formation (Upper Triassic) of Argentina. J. Vertebr. Paleontol. 13, 400–423 (1994).
Google Scholar
Langer, M. C. et al. Untangling the dinosaur family tree. Nature 551, E1–E3 (2017).
Google Scholar
Desojo, J. B. et al. The Late Triassic Ischigualasto Formation at Cerro Las Lajas (La Rioja, Argentina): fossil tetrapods, high-resolution chronostratigraphy, and faunal correlations. Sci. Rep. 10, 12782 (2020).
Google Scholar
Ree, R. H. & Smith, S. A. Maximum likelihood inference of geographic range evolution by dispersal, local extinction, and cladogenesis. Syst. Biol. 57, 4–14 (2008).
Google Scholar
Fitch, A. J., Lovelace, D. M. & Stocker, M. R. The oldest dinosaur from the northern hemisphere and the origins of Theropoda. in Program and Abstracts 80th Annual Meeting of the Society of Vertebrate Paleontology 140–141 (2020).
Evans, S. E. At the feet of the dinosaurs: the early history and radiation of lizards. Biol. Rev. 78, 513–551 (2003).
Google Scholar
Huttenlocker, A. K., Grossnickle, D. M., Kirkland, J. I., Schultz, J. A. & Luo, Z.-X. Late-surviving stem mammal links the lowermost Cretaceous of North America and Gondwana. Nature 558, 108–112 (2018).
Google Scholar
Lee, M. S. Y., Baron, M. G., Norman, D. B. & Barrett, P. M. Dynamic biogeographic models and dinosaur origins. Earth Environ. Sci. Trans. R. Soc. Edin. 109, 325–332 (2018).
Schaller, M. F., Wright, J. D. & Kent, D. V. A 30 Myr record of Late Triassic atmospheric pCO2 variation reflects a fundamental control of the carbon cycle by changes in continental weathering. GSA Bull. 127, 661–671 (2015).
Google Scholar
Baron, M. G., Norman, D. B. & Barrett, P. M. Baron et al. reply. Nature 551, E4–E5 (2017).
Google Scholar
Nesbitt, S. J. & Sues, H.-D. The osteology of the early-diverging dinosaur Daemonosaurus chauliodus (Archosauria: Dinosauria) from the Coelophysis Quarry (Triassic: Rhaetian) of New Mexico and its relationships to other early dinosaurs. Zool. J. Linnean Soc. 191, 0150–179 (2020).
Google Scholar
Baron, M. G. Pisanosaurus mertii and the Triassic ornithischian crisis: could phylogeny offer a solution? Hist. Biol. 31, 967–981 (2019).
Google Scholar
Benton, M. J. & Walker, A. D. Saltopus, a dinosauriform from the Upper Triassic of Scotland. Earth Environ. Sci. Trans. R. Soc. Edin. 101, 285–299 (2010).
Nesbitt, S. J., Barrett, P. M., Werning, S., Sidor, C. A. & Charig, A. J. The oldest dinosaur? A Middle Triassic dinosauriform from Tanzania. Biol. Lett. 9, 20120949 (2013).
Google Scholar
Agnolin, F. L. & Ezcurra, M. D. The validity of Lagosuchus talampayensis Romer, 1971 (Archosauria, Dinosauriformes), from the Late Triassic of Argentina. Breviora 565, 1–21 (2019).
Google Scholar
Goloboff, P. A., Farris, J. S. & Nixon, K. C. TNT, a free program for phylogenetic analysis. Cladistics 24, 774–786 (2008).
Google Scholar
Mesquite: a modular system for evolutionary analysis v.3.70 (2021).
Goloboff, P. A. & Szumik, C. A. Identifying unstable taxa: efficient implementation of triplet-based measures of stability, and comparison with Phyutility and RogueNaRok. Mol. Phylogenet. Evol. 88, 93–104 (2015).
Google Scholar
Wagner, P. J. A likelihood approach for evaluating estimates of phylogenetic relationships among fossil taxa. Paleobiology 24, 430–449 (1998).
Google Scholar
Kolaczkowski, B. & Thornton, J. W. Performance of maximum parsimony and likelihood phylogenetics when evolution is heterogeneous. Nature 431, 980–984 (2004).
Google Scholar
Goloboff, P. A., Pittman, M., Pol, D. & Xu, X. Morphological data sets fit a common mechanism much more poorly than DNA sequences and call into question the Mkv model. Syst. Biol. 68, 494–504 (2019).
Google Scholar
Goloboff, P. A., Torres, A. & Arias, J. S. Weighted parsimony outperforms other methods of phylogenetic inference under models appropriate for morphology. Cladistics 34, 407–437 (2018).
Google Scholar
Goloboff, P. A., Torres Galvis, A. & Arias, J. S. Parsimony and model-based phylogenetic methods for morphological data: comments on O’Reilly et al. Palaeontology 61, 625–630 (2018).
Google Scholar
Kammerer, C. F., Nesbitt, S. J., Flynn, J. J., Ranivoharimanana, L. & Wyss, A. R. A tiny ornithodiran archosaur from the Triassic of Madagascar and the role of miniaturization in dinosaur and pterosaur ancestry. Proc. Natl Acad. Sci. USA 117, 17932–17936 (2020).
Google Scholar
Ezcurra, M. D. et al. Enigmatic dinosaur precursors bridge the gap to the origin of Pterosauria. Nature 588, 445–449 (2020).
Google Scholar
Parry, L. A., Baron, M. G. & Vinther, J. Multiple optimality criteria support Ornithoscelida. R. Soc. Open Sci. 4, 170833 (2017).
Google Scholar
O’Reilly, J. E. et al. Bayesian methods outperform parsimony but at the expense of precision in the estimation of phylogeny from discrete morphological data. Biol. Lett. 12, 20160081 (2016).
Google Scholar
Koch, N. M., Garwood, R. J. & Parry, L. A. Fossils improve phylogenetic analyses of morphological characters. Proc. Royal Soc. B Biol. Sci. 288, 20210044 (2021).
Bouckaert, R. et al. BEAST 2.5: an advanced software platform for Bayesian evolutionary analysis. PLoS Comput. Biol. 15, e1006650 (2019).
Google Scholar
Drummond, A. J. & Suchard, M. A. Bayesian random local clocks, or one rate to rule them all. BMC Biol. 8, 114 (2010).
Google Scholar
Rambaut, A., Drummond, A. J., Xie, D., Baele, G. & Suchard, M. A. Posterior summarization in Bayesian phylogenetics using Tracer 1.7. Syst. Biol. 67, 901–904 (2018).
Google Scholar
Drummond, A. J. & Rambaut, A. BEAST: Bayesian evolutionary analysis by sampling trees. BMC Evol. Biol. 7, 214 (2007).
Google Scholar
Landis, M., Edwards, E. J. & Donoghue, M. J. Modeling phylogenetic biome shifts on a planet with a past. Syst. Biol. 70, 86–107 (2020).
Google Scholar
Landis, M. J., Freyman, W. A. & Baldwin, B. G. Retracing the Hawaiian silversword radiation despite phylogenetic, biogeographic, and paleogeographic uncertainty. Evolution 72, 2343–2359 (2018).
Google Scholar
R Core Team. R: A Language and Environment for Statistical Computing (R Foundation for Statistical Computing, 2020).
Matzke, N. J. Probabilistic historical biogeography: new models for founder-event speciation, imperfect detection, and fossils allow improved accuracy and model-testing. Front. Biogeogr. 5, 242–248 (2013).
Google Scholar
Matzke, N. J. Model selection in historical biogeography reveals that founder-event speciation is a crucial process in island clades. Syst. Biol. 63, 951–970 (2014).
Google Scholar
Ree, R. H. & Sanmartín, I. Conceptual and statistical problems with the DEC+J model of founder-event speciation and its comparison with DEC via model selection. J. Biogeogr. 45, 741–749 (2018).
Google Scholar
Aberhan, M. Bivalve palaeobiogeography and the Hispanic Corridor: time of opening and effectiveness of a proto-Atlantic seaway. Palaeogeogr. Palaeoclimatol. Palaeoecol. 165, 375–394 (2001).
Google Scholar
Schöllhorn, I. et al. Climate and environmental response to the break-up of Pangea during the Early Jurassic (Hettangian-Pliensbachian); the Dorset coast (UK) revisited. Glob. Planet. Change 185, 103096 (2020).
Google Scholar
Xiang, Y., Gubian, S., Suomela, B. & Hoeng, J. Generalized simulated annealing for global optimization: the GenSA package. R J. 5, 13–28 (2013).
Google Scholar
Revell, L. J. phytools: an R package for phylogenetic comparative biology (and other things). Methods Ecol. Evol. 3, 217–223 (2012).
Google Scholar
Apaldetti, C., Martínez, R. N., Cerda, I. A., Pol, D. & Alcober, O. An early trend towards gigantism in Triassic sauropodomorph dinosaurs. Nat. Ecol. Evol. 2, 1227–1232 (2018).
Google Scholar
Pol, D., Otero, A., Apaldetti, C. & Martínez, R. N. Triassic sauropodomorph dinosaurs from South America: the origin and diversification of dinosaur dominated herbivorous faunas. J. South Amer. Earth Sci. 107, 103145 (2021).
Google Scholar
Melo, T. P., Abdala, F. & Soares, M. B. The Malagasy cynodont Menadon besairiei (Cynodontia; Traversodontidae) in the Middle–Upper Triassic of Brazil. J. Vertebr. Paleontol. 35, e1002562 (2015).
Google Scholar
Parker, W. G. Revised phylogenetic analysis of the Aetosauria (Archosauria: Pseudosuchia); assessing the effects of incongruent morphological character sets. PeerJ 4, e1583 (2016).
Google Scholar
Parker, W. G. Redescription of Calyptosuchus (Stagonolepis) wellesi (Archosauria: Pseudosuchia: Aetosauria) from the Late Triassic of the Southwestern United States with a discussion of genera in vertebrate paleontology. PeerJ 6, e4291 (2018).
Google Scholar
Roberto-da-Silva, L. et al. A new aetosaur from the Upper Triassic of the Santa Maria Formation, southern Brazil. Zootaxa 3764, 240–278 (2014).
Google Scholar
Alcober, O. A. & Martinez, R. N. A new herrerasaurid (Dinosauria, Saurischia) from the Upper Triassic Ischigualasto Formation of northwestern Argentina. Zookeys 19, 55–81 (2010).
Brinkman, D. B. & Sues, H.-D. A staurikosaurid dinosaur from the Upper Triassic Ischigualasto Formation of Argentina and the relationships of the Staurikosauridae. Palaeontology 30, 493–503 (1987).
Pacheco, C. et al. Gnathovorax cabreirai: a new early dinosaur and the origin and initial radiation of predatory dinosaurs. PeerJ 7, e7963 (2019).
Google Scholar
Angielczyk, K. D., Hancox, P. J. & Nabavizadeh, A. A redescription of the Triassic kannemeyeriiform dicynodont Sangusaurus (Therapsida, Anomodontia), with an analysis of its feeding system. J. Vertebr. Paleontol. 37, 189–227 (2017).
Google Scholar
Langer, M. C., da Rosa, Á. A. S. & Montefeltro, F. C. Supradapedon revisited: geological explorations in the Triassic of southern Tanzania. PeerJ 5, e4038 (2017).
Google Scholar
Montefeltro, F. C., Langer, M. C. & Schultz, C. L. Cranial anatomy of a new genus of hyperodapedontine rhynchosaur (Diapsida, Archosauromorpha) from the Upper Triassic of southern Brazil. Earth Environ. Sci. Trans. R. Soc. Edin. 101, 27–52 (2010).